Pupina artata Benson, 1856
“It has some affinity with the Australian species of the genus. Although the following shell is not Burmese, yet as it is found in the Ultra-Gangetic portions of the Bengal Provinces, which are geographically a continuation of the same zoological tract, I shall give it a place here.” (Benson, 1856)
“My friend Mr. Theobald has, I fear, led Mr. Hanley into an error in the Con. Indica regarding this group; the form from Thyet Myo called P. blanfordiana by Mr. Theobald is exceedingly close to P. artata. I consider it nothing but a mere local variety; the shell figured for it in the Con. Indica is quite different, being exceedingly close to P. arula (so close that I am inclined to doubt its being more than a variety); the figure is an exact representation of the type of P. peguensis. A comparison with Benson's admirable description, where he points out its close affinities to his P. arula, should convince even my friend, Mr. Theobald himself. In any case P. blanfordiana must be altogether eliminated from the list of species, if my view is correct, as it is a mere local form of P. artata; if the figure, however, in the Con. Indica really represents Theobald's form, it becomes a synonym of P. peguensis.” (Nevill, 1878)
“J'ai rencontré cette espèce dans la jungle basse aux environs de Lahat, Ipoh, Gôping, etc. (Kinta), elle vit sur les détritus végétaux et est assez rare.” (Morgan, 1885)
“The dead, worn example from Chieng Dao was 6.34 mm. high, diameter 4.38 mm., with 5 5/8 whorls. A live collected shell from Doi Sutep was 6.21 mm. high, diameter 4.71 mm., with slightly less than 5 3/8 whorls.” (Solem, 1966)
Benson (1856) original descriptions on Pupina artata – “Testa imperforata, pupiformi, ovato-acuminata, politissima, fuscocornea, translucente, vel hyalina; spira ovato-conoidea, apice obtusiusculo, sutura callosa, lineari; anfractibus 5½ convexiusculis, ultimo 1/3 testae partem superante; apertura verticali, circulari, bicanaliculata; peristomate obtusiusculo, margine parietali superne linguam acutam, callo verticali elongato, cum margine dextro subparallelo, marginatum, exhibente, infra cum basali canalem incisum, ascendentem, extus calloso-marginatum, efformante. Operculo testaceo pauci-spirali, concaviusculo, sutura elevata.”
Kobelt (1902) descriptions on Pupina (Tylotoechus) artata – “Schale ungenabelt, spitz eiförmig, sehr glatt, durchscheinend, hornbraun; Apex ziemlich spitz; Naht schwielig, linienförmig; 5½ ziemlich gewölbte Windungen, letzte 1/3 der Höhe einnehmend; Mündung senkrecht, kreisrund, mit 2 Kanälen; Mundrand ziemlich stumpf, an der vorletzten Windung oben ein zungenfönniger Fortsatz, der durch einen langen, senkrechten, dem recliten Rande parallelen Callus begrenzt wird, unten mit dem Unterrand einen eingeschnittenen, ansteigenden, aussen schwielig begrenzten Kanal bildend. Deckel mit wenigen Windungen und erhobener Naht, leicht konkav.”
“Foot oblong, the sole being somewhat truncate in front and slightly angled at each side anteriorly, hinder extremity narrowed and pointed; muzzle declivous, entire; tentacula short, subulate, and swollen all round at the base; eyes black and prominent, situated on the hinder and external part of the basal swelling. The operculum is rather thick, horny, rounded at the thickened edge, and consists of 4½-5 concave whorls divided by a raised edge. The inner surface has the umbonal region a little elevated. In my first description of the shell I stated that the operculum was calcareous, with few whorls. Dr. Pfeiffer, who had the specimen before him, made no alteration in the description, but observed that my characters were abnormal. The paucity of whorls was intended to be comparative with reference to the allied genus Megalomastoma. On taking out the operculum of that specimen, its substance appears evidently to be horny; but neither in this species, nor in the Khasia P. imbricifera, which has a still thicker horny operculum, with a very prominent umbo internally, can it be said to answer Pfeiffer's generic description of " membranaceous." The animal was easily revived in two specimens with the operculum received from Capt. Haughton, and began at once to move about freely and fearlessly. No organ corresponding with the slits at the upper or lower part of the aperture can be detected.” (Benson, 1859)
Pupina artata – “Long. 6, diam. 3½ mill.” (Benson, 1856); Pupina (Tylotoechus) artata – “H. 7, Durchm. 4. Mündungsweite 3.75 mm.” (Kobelt, 1902)
Type locality – “ad Moulmein satis frequens” (Benson, 1856)
Other localities – “Lahat, Ipoh, Gôping, etc. (Kinta)” (Morgan, 1885); “Foot of a hill near the hot springs, near Tandjong Rambutan, NE. of Ipoh, Perak” leg. Benthem-Jutting/1946 (Benthem-Jutting, 1960); “Thailand; Chieng Dao at 1,100 meters” (Solem, 1966); “Thailand; Doi Sutep at 1,100 meters” (Solem, 1966); “limestone island C, Temenggor dam, 230 m” leg. Davison/Aug. 28, 1993 (Davison, 1995)